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Lepraria bergensis Tønsberg |
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Nomenclatural data
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Graphis Scripta 14: 47 (2002); type: Norway. Hordaland: Bergen, Haukeland/Landaås, Vognstølen, 60°22.2'N, 5°21.8'E, 25 Nov. 2000, Tønsberg 28875 (BG—holotypus; B, H, NMW—isotypi).
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Morphology
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Thallus crustose, leprose, with membranous appearance; pale bluish grey; relatively thin, up to 220 µm excluding loose soredia, relatively soft, not very firmly attached to the substrate; young thalli form small rosettes up to few mm in diam., later fused patches 1 cm or more in diam. form; margin usually delimited, sharply raised (in saxicolous specimens), lobes usually distinct, up to few mm wide; cortex absent; medulla distinct in places, white; hypothallus not evident or orange-brown to brown, never conspicuous and not visible from above, hyphae below thallus projecting, orange-brown to brown or colourless, lower surface present, white (at least along margin), pale yellowish or brown; prothallus absent; areoles absent; squamules absent; soredia abundant, very fine, up to 25 µm in diam., mostly loosely packed; projecting hyphae present or absent, short; consoredia abundant, up to 200(-340) µm; isidia-like structures absent. Photobiont trebouxioid, to 16 µm in diameter.
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Chemistry
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Atranorin, rangiformic acid, norrangiformic acid (trace), fragilin, 7-chloroemodin, emodin, parietin, A01-anthrone; the anthraquinones are located mostly in the subthalline hyphae. K+ faint yellow, subthalline hyphae K+ purple (in patches), C–, Pd–.
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Remarks
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The morphologically similar species include L. bergensis, L. membranacea, L. normandinoides, L. sipmaniana that all have different diagnostic substances. L. bergensis produces atranorin, rangiformic acid and anthraquinones. L. membranacea always contains pannaric acid. L. normandinoides usually produces protocetraric acid and/or fumarprotocetraric acid besides constant roccellic acid and atranorin. L. sipmaniana is distinguished by pannaric acid 6-methylester and usually also anthraquinones. L. humida, too, produces anhraquinones, but lacks lobes. Several additional taxa can develop well developed lobes, see the discussion under L. membranacea.
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Ecology and distribution
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Substrate and ecology: on siliceous rock, also on mosses on rock, shaded surfaces. Distribution: Europe – Norway, Great Britain, Germany (Spribille & Tønsberg 2007).
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