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Lepraria adhaerens K. Knudsen, Elix & Lendemer |
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Nomenclatural data
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Opuscula Philolichenum, 4: 5 (2007); type: U.S.A. California, San Diego Co., Torrey Pines State Park, 32° 54' 58"N, 117° 19' 56"W, 13. Apr. 2005, K. Knudsen et al. 2700 (UCR—holotypus; ASU, CANB, PH, SD, UGDA—isotypi).
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Morphology
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Thallus crustose, leprose, with granular appearance; light blue-grey, sometimes with a dull brownish tinge; thin to thick (up to 1 mm), relatively hard; shape irregular, individual colonies up to 2 cm, fused patches up to several meters; margin diffuse, lobes absent; cortex absent; medulla absent, but older thalli forming a lower necral layer of gelatinized granules; lower surface absent; areoles present in well developed thalli, divided by deep fissures; squamules absent; thallus surfaces without soredia absent, soredia abundant, fine to medium, 40-100 µm in diam., densely packed; wall present; projecting hyphae absent, but frequently with thin colourless hyphae acting as anchors or rhizines; soredia clumped together; isidia-like structures absent. Photobiont chlorococcoid, 10-14 µm in diam., often aggregating.
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Chemistry
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Pannarin and zeorin (major to trace, very rarely absent), accessories include norpannarin ± (trace), dechloropannarin ± (trace), hypopannarin ± (trace), atranorin ± (trace). K–, C–, KC–, Pd+ orange.
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Remarks
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L. santamonicae and L. coriensis produce chlorodepsidones argopsin and norargopsin that are biosequentially related to pannarin, but otherwise these species differ chemically and morphologically. The morphology of L. adhaerens is most similar to that of L. santamonicae (Knudsen & Elix 2007), but is not radically different from other unstratified Lepraria species with indeterminate thalli which lack lobes. The most characteristic feature of L. adhaerens is the small attaching hyphae and the tendency of granules to adhere to one another and the substrate.
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Ecology and distribution
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Substrate and ecology: mosses and lichens on different types of rock, rarely directly on soil or rock; open habitats, also those exposed to rain and sunlight, but sheltered places in snowy areas. Distribution: North America.
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