For a detailed description, see
Clerc (2004b, 2007). Thallus shrubby to subpendent, richly branched; branching mainly isotomic-dichotomous; base blackened, often with transverse annular cracks; fibrils present already on very young branches, acting like diaspores – fibrils are shed leaving small fibercles, while soredia and isidiomorphs may develop inside these fibercles; papillae present, predominantly on main branches; soralia minute, plane, later becoming confluent; isdiomorphs short, numerous on young soralia but present also on mature ones. Cortex rather thick; medulla thin and dense.